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  1. Article ; Online: Systematic revision of Stegodera Martens, 1876 (Gastropoda, Stylommatophora, Camaenidae), with description of a new genus

    Zhe-Yu Chen / Zhi-Tong Lyu / Min Wu

    ZooKeys, Vol 1059, Iss , Pp 1-

    2021  Volume 21

    Abstract: The monotypic genus Stegodera Martens, 1876 is systematically revised based on anatomical and morphological examination of freshly collected specimens. A new species from southern Hunan, which resembles Stegodera angusticollis, is confirmed to represent ... ...

    Abstract The monotypic genus Stegodera Martens, 1876 is systematically revised based on anatomical and morphological examination of freshly collected specimens. A new species from southern Hunan, which resembles Stegodera angusticollis, is confirmed to represent a new genus evidenced by comparative shell morphology and anatomy as well as by molecular phylogenetic analyses. The new genus might be more closely related to Stegodera and Nesiohelix Kuroda & Emura, but differs anatomically from the latter two genera by the absence of a dart apparatus.
    Keywords Zoology ; QL1-991
    Language English
    Publishing date 2021-09-01T00:00:00Z
    Publisher Pensoft Publishers
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  2. Article ; Online: A new species of the genus Lycodon (Serpentes, Colubridae) from Guangxi, China

    Jian Wang / Shuo Qi / Zhi-Tong Lyu / Zhao-Chi Zeng / Ying-Yong Wang

    ZooKeys, Vol 954, Iss , Pp 85-

    2020  Volume 108

    Abstract: A new species of colubrid snake, Lycodon cathaya sp. nov., is described based on two adult male specimens collected from Huaping Nature Reserve, Guangxi, southern China. In a phylogenetic analyses, the new species is shown to be a sister taxon to the ... ...

    Abstract A new species of colubrid snake, Lycodon cathaya sp. nov., is described based on two adult male specimens collected from Huaping Nature Reserve, Guangxi, southern China. In a phylogenetic analyses, the new species is shown to be a sister taxon to the clade composed of L. futsingensis and L. namdongensis with low statistical support, and can be distinguished from all known congeners by the significant genetic divergence in the mitochondrial cytochrome b gene fragment (p-distance ≥ 7.9%), and morphologically by the following combination of characters: (1) dorsal scales in 17–17–15 rows, smooth throughout; (2) supralabials eight, third to fifth in contact with eye, infralabials nine; (3) ventral scales 199–200 (plus two preventral scales), subcaudals 78; (4) loreal single, elongated, in contact with eye or not, not in contact with internasals; (5) a single preocular not in contact with frontal, supraocular in contact with prefrontal, two postoculars; (6) maxillary teeth 10 (4+2+2+2); (7) two anterior temporals, three posterior temporals; (8) precloacal plate entire; (9) ground color from head to tail brownish black, with 31–35 dusty rose bands on body trunk, 13–16 on tail; (10) bands in 1–2 vertebral scales broad in minimum width; (11) bands separate ground color into brownish black ellipse patches arranged in a row along the top of body and tail; (12) elliptical patches in 3–6 scales of the vertebral row in maximum width; (13) ventral surface of body with wide brownish black strip, margined with a pair of continuous narrow greyish white ventrolateral lines. With the description of the new species, 64 congeners are currently known in the genus Lycodon, with 16 species occurring in China.
    Keywords Zoology ; QL1-991
    Subject code 590
    Language English
    Publishing date 2020-07-01T00:00:00Z
    Publisher Pensoft Publishers
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  3. Article ; Online: Generic classification of Asian horned toads (Anura

    Zhi-Tong Lyu / Shuo Qi / Jian Wang / Si-Yu Zhang / Jian Zhao / Zhao-Chi Zeng / Han Wan / Jian-Huan Yang / Yun-Ming Mo / Ying-Yong Wang

    Zoological Research, Vol 44, Iss 2, Pp 380-

    Megophryidae: Megophryinae) and monograph of Chinese species

    2023  Volume 450

    Abstract: The subfamily Megophryinae, as a representative batrachian group of the Oriental Realm and one of the most diverse groups of amphibians, has attracted considerable attention due to continued conjecture regarding its generic classification and failure to ... ...

    Abstract The subfamily Megophryinae, as a representative batrachian group of the Oriental Realm and one of the most diverse groups of amphibians, has attracted considerable attention due to continued conjecture regarding its generic classification and failure to reach a satisfactory consensus. China boasts the richest diversity of Asian horned toads, containing some two thirds of the total species cataloged. However, most species have a complicated taxonomic history, resulting in multiple misidentifications. As such, an overall clarification of historical records and regional checklists is required. In the current investigation, we established the phylogeny of the Asian horned toads and performed detailed examinations with redefinitions of several important morphological traits. Based on the phylogenetic relationships and morphological differences, we propose a new ten-genus classification for the Asian horned toad subfamily Megophryinae: i.e., Brachytarsophrys, Atympanophrys, Grillitschia, Sarawakiphrys gen. nov., Jingophrys gen. nov., Xenophrys, Megophrys, Pelobatrachus, Ophryophryne, and Boulenophrys. Revisions on the diagnosability, distribution, and content of each genus are provided. Furthermore, we present a careful review of the taxonomic history of Asian horned toad species from China and provide a monograph of congeners, including six species of Brachytarsophrys, four species of Atympanophrys, five species of Jingophrys gen. nov., 10 species of Xenophrys, two species of Ophryophryne, and 60 species of Boulenophrys. Finally, we discuss the importance of traditional morphological traits based on multiple populations in taxonomic work as well as taxonomic inflation caused by the genetic species delimitation.
    Keywords atympanophrys ; boulenophrys ; megophrys ; jingophrys gen. nov ; review ; sarawakiphrys gen. nov ; taxonomy ; xenophrys ; Zoology ; QL1-991
    Subject code 590 ; 580
    Language English
    Publishing date 2023-03-01T00:00:00Z
    Publisher Science Press, PR China
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  4. Article ; Online: A new species of Music frog (Anura, Ranidae, Nidirana) from Mt Daming, Guangxi, China

    Zhi-Tong Lyu / Zhong Huang / Xiao-Wen Liao / Li Lin / Yong Huang / Ying-Yong Wang / Yun-Ming Mo

    ZooKeys, Vol 1059, Iss , Pp 35-

    2021  Volume 56

    Abstract: Nidirana guangxiensis sp. nov., a new music frog species, is proposed, based on a series of specimens collected from Mt Daming, Guangxi, southern China. The new species is close to N. yeae, N. daunchina, N. yaoica, and N. chapaensis from southwestern and ...

    Abstract Nidirana guangxiensis sp. nov., a new music frog species, is proposed, based on a series of specimens collected from Mt Daming, Guangxi, southern China. The new species is close to N. yeae, N. daunchina, N. yaoica, and N. chapaensis from southwestern and south-central China and northern Indochina, while the relationships among these species remain unresolved. Nidirana guangxiensis sp. nov. can be distinguished from all known congeners by the genetic divergences in the mitochondrial 16S and COI genes, the behavior of nest construction, the advertisement call containing 6–11 rapidly repeated regular notes, and a combination of morphological characteristics. Furthermore, the Nidirana populations recorded in Guangxi are clarified in this work, providing valuable new information on the knowledge of the genus Nidirana.
    Keywords Zoology ; QL1-991
    Language English
    Publishing date 2021-09-01T00:00:00Z
    Publisher Pensoft Publishers
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  5. Article ; Online: Taxonomic revision of Amolops chunganensis (Pope, 1929) (Amphibia

    Ke Jiang / Jin-Long Ren / Zhi-Tong Lyu / Dan Wang / Zeng Wang / Ke Lv / Jia-Wei Wu / Jia-Tang Li

    Zoological Research, Vol 42, Iss 5, Pp 574-

    Anura) and description of a new species from southwestern China, with discussion on Amolops monticola group and assignment of species groups of the genus Amolops

    2021  Volume 591

    Abstract: Amolops chunganensis is a species complex and reported widely from eastern, southern, and southwestern China. Based on molecular data of 19 populations of A. chunganensis sensu lato from China, including the population from Mt. Wuyi (type locality), we ... ...

    Abstract Amolops chunganensis is a species complex and reported widely from eastern, southern, and southwestern China. Based on molecular data of 19 populations of A. chunganensis sensu lato from China, including the population from Mt. Wuyi (type locality), we recognize A. chunganensis sensu stricto and provide an expanded description based on the topotypic specimens. Combining morphological and molecular data, we describe a new species, Amolops chaochin sp. nov., from southwestern China, which was previously identified as A. chunganensis. The new species is distinguished from all other species in the A. monticola group by: (1) moderate body size, SVL 35.3−39.2 mm in males (n=7), and 50.5−54.4 mm in females (n=7); (2) distinct tympanum, larger than half of eye diameter; (3) small tooth-like projection on anteromedial edge of mandible; (4) circummarginal groove on all fingers; (5) white tubercles on dorsal side of posterior body in both sexes; (6) distinct tubercles on dorsal thigh and white spinose tubercles on dorsal tibia in both sexes; (7) white tubercles on posterior region of tympanum in males; (8) toe webbing reaching disk by dermal fringe on inner side of toe II; (9) vomerine teeth present; (10) transverse bands on dorsal limbs; (11) external vocal sacs present in males. We further reviewed the assignment of Amolops groups, with an overall revision of membership and diagnosis of all species groups.
    Keywords amolops chaochin sp. nov ; complex ; distribution ; amolops gerbillus ; amolops chayuensis group ; amolops spinapectoralis group ; Zoology ; QL1-991
    Subject code 590
    Language English
    Publishing date 2021-09-01T00:00:00Z
    Publisher Science Press, PR China
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  6. Article ; Online: A new species of the genus Takydromus (Squamata, Lacertidae) from southwestern Guangdong, China

    Jian Wang / Zhi-Tong Lyu / Chen-Yu Yang / Yu-Long Li / Ying-Yong Wang

    ZooKeys, Vol 871, Iss , Pp 119-

    2019  Volume 139

    Abstract: A new species, Takydromus yunkaiensis J. Wang, Lyu, & Y.Y. Wang, sp. nov. is described based on a series of specimens collected from the Yunkaishan Nature Reserve located in the southern Yunkai Mountains, western Guangdong Province, China. The new ... ...

    Abstract A new species, Takydromus yunkaiensis J. Wang, Lyu, & Y.Y. Wang, sp. nov. is described based on a series of specimens collected from the Yunkaishan Nature Reserve located in the southern Yunkai Mountains, western Guangdong Province, China. The new species is a sister taxon to T. intermedius with a genetic divergence of 8.0–8.5% in the mitochondrial cytochrome b gene, and differs from all known congeners by a combination of the following morphological characters: (1) body size moderate, SVL 37.8–56.0 mm in males, 42.6–60.8 mm in females; (2) dorsal ground color brown; ventral surface green to yellow-green, but light blue-green on chin and throat, posteriorly green in adult males; (3) dorsolateral lines paired, strikingly yellowish-white bordered by black above and below, invisible or indistinct in juveniles and adult females; (4) flanks of body blackish brown with light brown marks in adult males; (5) presence of four pairs of chin-shields; (6) four supraoculars on each side; (7) presence of a row of supracilary granules that separate supracilaries from supraoculars; (8) two postnasals; (9) enlarged dorsal scales in six longitudinal rows on trunk of body, with strong keel; (10) enlarged ventral scales in six longitudinal rows, strongly keeled in males, smooth but outermost rows weakly keeled in females; (11) enlarged and keeled lateral scales in a row above ventrals; (12) femoral pores 2–3 on each side; (13) subdigital lamellae 20–23 under the fourth finger, 23–30 under the fourth toe; and (14) the first 2–3 subdigital lamellae under the fourth toe divided. The discovery of Takydromus yunkaiensis sp. nov. brings the total number of species of this genus to 24, of which nine occur in mainland China.
    Keywords Zoology ; QL1-991
    Subject code 590
    Language English
    Publishing date 2019-08-01T00:00:00Z
    Publisher Pensoft Publishers
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  7. Article ; Online: A new Leptobrachella species (Anura, Megophryidae) from South China, with comments on the taxonomic status of L. chishuiensis and L. purpurus

    Jian Wang / Shuo Qi / Ke-Yuan Dai / Zhi-Tong Lyu / Zhao-Chi Zeng / Hong-Hui Chen / Yuan-Qiu Li / Yong-You Zhao / Yun-Ze Wang / Ying-Yong Wang

    Zoosystematics and Evolution, Vol 98, Iss 1, Pp 165-

    2022  Volume 180

    Abstract: A new species of Leaf Litter Toad, Leptobrachella shimentaina sp. nov., is described from the Shimentai and Luokeng nature reserves of northern Guangdong Province, southern China. The new taxon can be distinguished from all recognized congeners by a ... ...

    Abstract A new species of Leaf Litter Toad, Leptobrachella shimentaina sp. nov., is described from the Shimentai and Luokeng nature reserves of northern Guangdong Province, southern China. The new taxon can be distinguished from all recognized congeners by a combination of discrete morphological character state differences relating to its small body size (SVL 26.4–28.9 mm in six adult males, 30.1 and 30.7 mm in two adult females); a number of apparently fixed color pattern character differences (including eye coloration and color pattern features from dorsal, ventral, and dorsolateral surfaces of its head, body, limbs, and ventrum); the morphological and discrete characteristics of the external phenotype (the skin texture of dorsum and ventrum, the presence of supra-axillary and ventrolateral glands, the wide dermal fringes and rudimentary webbing on toes, and the uninterrupted longitudinal ridges under toes). Two samples of this new species previously were proposed as representing a new, unnamed species. We now substantiate this claim by providing diagnostic comparisons of discrete character differences. In addition, we also discuss taxonomic uncertainty surrounding the identity of two congeners, L. chishuiensis and L. purpurus, which we interpret as indicative of taxonomic inflation in the species-rich subfamily Megophryidae.
    Keywords Biology (General) ; QH301-705.5
    Subject code 590
    Language English
    Publishing date 2022-06-01T00:00:00Z
    Publisher Pensoft Publishers
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  8. Article ; Online: Taxonomic clarifications on the floating frogs (Anura

    Zhi-Tong Lyu / Jian Wang / Zhao-Chi Zeng / Lin Luo / Yan-Wu Zhang / Chun-Peng Guo / Jin-Long Ren / Shuo Qi / Yun-Ming Mo / Ying-Yong Wang

    Vertebrate Zoology, Vol 72, Iss , Pp 495-

    Dicroglossidae: Occidozyga sensu lato) in southeastern China

    2022  Volume 512

    Abstract: The recognition for the floating frogs’ genus Occidozyga is in controversy for decades, and the species diversity of these frogs has recently been considered to be underestimated. In southeastern China, two floating frog species are currently recorded, ... ...

    Abstract The recognition for the floating frogs’ genus Occidozyga is in controversy for decades, and the species diversity of these frogs has recently been considered to be underestimated. In southeastern China, two floating frog species are currently recorded, namely Occidozyga lima and Occidozyga martensii. However, their current taxonomic statuses are unresolved after a series of recent taxonomic revisions. In this work, we perform morphological examinations and phylogenetic analyses on these two recorded floating frogs from southeastern China, to clarify their current taxonomic placements. The population previously recorded as Occidozyga lima should be re-assigned to the nomenclature Occidozyga obscura comb. nov., and the population previously recorded as Occidozyga martensii should be an undescribed species which is erected as Occidozyga lingnanica sp. nov. in this work.
    Keywords Zoology ; QL1-991
    Subject code 590
    Language English
    Publishing date 2022-07-01T00:00:00Z
    Publisher Pensoft
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  9. Article ; Online: A definition of the Goniurosaurus yingdeensis group (Squamata, Eublepharidae) with the description of a new species

    Shuo Qi / L. Lee Grismer / Zhi-Tong Lyu / Liang Zhang / Pi-Peng Li / Ying-Yong Wang

    ZooKeys, Vol 986, Iss , Pp 127-

    2020  Volume 155

    Abstract: A definition of the Goniurosaurus yingdeensis group is presented in this study, on the basis of morphological and phylogenetic analyses based on a series of additional specimens. Moreover, a new species of this group, Goniurosaurus varius sp. nov., is ... ...

    Abstract A definition of the Goniurosaurus yingdeensis group is presented in this study, on the basis of morphological and phylogenetic analyses based on a series of additional specimens. Moreover, a new species of this group, Goniurosaurus varius sp. nov., is proposed for northern Guangdong Province, China. The new species can be distinguished from the other two congeners of this group by the following unique characters: one or two internasals; enlarged supraorbital tubercles absent; paravertebral tubercles between limb insertions 27–29; dorsal tubercle rows at midbody 21–24; ten precloacal pores in males and absent in females; body bands with black spots; iris orange-red.
    Keywords Zoology ; QL1-991
    Language English
    Publishing date 2020-11-01T00:00:00Z
    Publisher Pensoft Publishers
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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  10. Article ; Online: Four new species of Asian horned toads (Anura, Megophryidae, Megophrys) from southern China

    Zhi-Tong Lyu / Yuan-Qiu Li / Zhao-Chi Zeng / Jian Zhao / Zu-Yao Liu / Guo-Xin Guo / Ying-Yong Wang

    ZooKeys, Vol 942, Iss , Pp 105-

    2020  Volume 140

    Abstract: Recent phylogenetic analysis encompassing multilocus nuclear-gene and matrilineal mtDNA genealogy has revealed a series of cryptic species of the subgenus Panophrys within genus Megophrys from southern and eastern China. This study demonstrates that the ... ...

    Abstract Recent phylogenetic analysis encompassing multilocus nuclear-gene and matrilineal mtDNA genealogy has revealed a series of cryptic species of the subgenus Panophrys within genus Megophrys from southern and eastern China. This study demonstrates that the Panophrys specimens from the hilly areas among Guangdong, Guangxi and Hunan can be morphologically distinguished from all recognized congeners, thereby providing additional supports for the recognitions of four new species of Panophrys, namely Megophrys (Panophrys) mirabilis Lyu, Wang & Zhao, sp. nov. from northeastern Guangxi, Megophrys (Panophrys) shimentaina Lyu, Liu & Wang, sp. nov. from northern Guangdong, and Megophrys (Panophrys) xiangnanensis Lyu, Zeng & Wang, sp. nov. and Megophrys (Panophrys) yangmingensis Lyu, Zeng & Wang, sp. nov. from southern Hunan. The descriptions of these species take the number of Megophrys species to 101, 46 of which belong to the subgenus Panophrys.
    Keywords Zoology ; QL1-991
    Subject code 590
    Language English
    Publishing date 2020-06-01T00:00:00Z
    Publisher Pensoft Publishers
    Document type Article ; Online
    Database BASE - Bielefeld Academic Search Engine (life sciences selection)

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